diff --git a/src/core/inlines.js b/src/core/inlines.js
index 2b627582c1..71f8308b98 100644
--- a/src/core/inlines.js
+++ b/src/core/inlines.js
@@ -65,7 +65,7 @@ const l10n = getIntlData(localizationStrings);
 // add support.
 const inlineCodeRegExp = /(?:`[^`]+`)(?!`)/; // `code`
 const inlineIdlReference = /(?:{{[^}]+\?*}})/; // {{ WebIDLThing }}, {{ WebIDLThing? }}
-const inlineVariable = /\B\|\w[\w\s]*(?:\s*:[\w\s&;"<>]+\??)?\|\B/; // |var : Type?|
+const inlineVariable = /\B\|\w[\w\s]*(?:\s*:[\w\s&;"?<>]+\??)?\|\B/; // |var : Type?|
 const inlineCitation = /(?:\[\[(?:!|\\|\?)?[\w.-]+(?:|[^\]]+)?\]\])/; // [[citation]]
 const inlineExpansion = /(?:\[\[\[(?:!|\\|\?)?#?[\w-.]+\]\]\])/; // [[[expand]]]
 const inlineAnchor = /(?:\[=[^=]+=\])/; // Inline [= For/link =]
diff --git a/tests/spec/core/inlines-spec.js b/tests/spec/core/inlines-spec.js
index 96e5654e0f..7e26c43818 100644
--- a/tests/spec/core/inlines-spec.js
+++ b/tests/spec/core/inlines-spec.js
@@ -171,7 +171,7 @@ describe("Core - Inlines", () => {
         <p id="h"> TEXT |var: Generic&lt;int&gt;| TEXT |var2: Generic&lt;unsigned short int&gt;| </p>
       </section>
       <section>
-        <p id="nulls"> |var 1: null type spaces?| and |var 2 : NullableType?| </p>
+        <p id="nulls"> |var 1: null type spaces?| and |var 2 : NullableType?| and |var 3: Generic&lt;NullableType?&gt;|</p>
       </section>
     `;
     const doc = await makeRSDoc(makeStandardOps(null, body));
@@ -223,11 +223,14 @@ describe("Core - Inlines", () => {
     expect(h[1].textContent).toBe("var2");
     expect(h[1].dataset.type).toBe("Generic<unsigned short int>");
 
-    const [nullVar1, nullVar2] = doc.querySelectorAll("#nulls > var");
-    expect(nullVar1.textContent).toBe("var 1");
-    expect(nullVar1.dataset.type).toBe("null type spaces?");
-    expect(nullVar2.textContent).toBe("var 2");
-    expect(nullVar2.dataset.type).toBe("NullableType?");
+    const nulls = doc.querySelectorAll("#nulls > var");
+    expect(nulls).toHaveSize(3);
+    expect(nulls[0].textContent).toBe("var 1");
+    expect(nulls[0].dataset.type).toBe("null type spaces?");
+    expect(nulls[1].textContent).toBe("var 2");
+    expect(nulls[1].dataset.type).toBe("NullableType?");
+    expect(nulls[2].textContent).toBe("var 3");
+    expect(nulls[2].dataset.type).toBe("Generic<NullableType?>");
   });
 
   it("expands inline references and they get classified as normative/informative correctly", async () => {